Discussion of imalsogreg's comment: So much modeling and extrapolation about vision comes from anesthetized V1 recordings (or recordings in awake paralyzed cats). Recordings in awake, closer-to-freely-moving (treadmill) mice would have seemed like a sanity check, to make sure that the old rules hold in a brain that's active. The amount of V1 activity that has nothing to do with vision at all suggests that the old rules are an oversimplification. "Primary visual cortex" could actually be a misnomer - this study should encourage us to go back and record the whole visual pathway in moving animals.